Mgr. Adam Tomašových, PhD.

scientist, head of Department of Paleobiology and Organismal Evolution

Workplace:Bratislava, Dúbravská cesta 9
Personal webpage:



Research interests:

  • Quality of the fossil record, fossilization dynamic and temporal and spatial scaling of fossil assemblages: empirical and modeling approaches
  • Biogeography, macroecology and community ecology of marine invertebrate communities, especially with mollusks and brachiopods
  • Evolutionary ecology of Mesozoic-Cenozoic marine invertebrate ecosystems and End-Triassic mass extinction event: paleoecology and geochemistry
  • Conservation paleobiology of marine ecosystems


Professional preparation and appointments:

2002 Mgr. (M.S.) Comenius University, Bratislava, Slovakia (Geology)

2006 Ph.D., Würzburg University, Germany (Geology)

2006-2011 – University of Chicago, Department of Geophysical Sciences, Research Associate – Postdoctoral Fellow

2011-present – Earth Science Institute of Slovak Academy of Sciences, Bratislava, Researcher


Web Of Science publications:

[57] BAJNAI, Dávid – FIEBIG, J. – TOMAŠOVÝCH, Adam – GARCIA, Sara Milner – ROLLION-BARD, Claire – RADDATZ, J. – LÖŢ ER, Niklas – PRIMO-RAMOS, Cristina – BRAND, Uwe. Assessing kinetic fractionation in brachiopod calcite using clumped isotopes. In Scientific Reports, 2018, vol. 8, art. no. 533.

[56] KOŠŤÁK, Martin – SCHLÖGL, Ján – CULKA, Adam – TOMAŠOVÝCH, Adam – MAZUCH, Martin. The unique preservation of Sepia soft tissues in the Miocene deposits (Serravalian, Vienna Basin): Implications for the origin of microbodies in the fossil record. In Palaeogeography, Palaeoclimatology, Palaeoecology, 2018, vol. 493, p. 111-118.

[55] MAUTNER, Anna-Katharina – GALLMETZER, Ivo – HASELMAIR, Alexandra – SCHNEDL, Sara-Maria – TOMAŠOVÝCH, Adam – ZUSCHIN, Martin. Holocene ecosystem shifts and human-induced loss of Arca and Ostrea shell beds in the north-eastern Adriatic Sea. In Marine Pollution Bulletin, 2018, vol. 126, p. 19-30.

[54] BIZJACK, Matthew T. – KIDWELL, Susan M. – VELARDE, Ronald G. – LEONARD-PINGEL, Jill – TOMAŠOVÝCH, Adam. Detecting, sourcing, and age-dating dredged sediments on the open shelf, southern California, using dead mollusk shells. In Marine Pollution Bulletin, 2017, vol. 114, no. 1, p. 448-465.

[53] GALLMETZER, Ivo – HASELMAIR, Alexandra – TOMAŠOVÝCH, Adam – STACHOWITSCH, Michael – ZUSCHIN, Martin. Responses of molluscan communities to centuries of human impact in the northern Adriatic Sea. In PLoS ONE, 2017, vol.12., no. 7, 32 p.

[52] TOMAŠOVÝCH, Adam – KIDWELL, Susan M. Nineteenth-century collapse of a benthic marine ecosystem on the open continental shelf. In Proceedings of the Royal Society : B – Biological Sciences, 2017, vol. 284, no. 1856, 9 p.

[51] TOMAŠOVÝCH, Adam – GALIMETZER, Ivo – HASELMAIR, Alexandra – KAUFMAN, D. S. – VIDOVIĆ, Jelena – ZUSCHIN, Martin. Stratigraphic unmixing reveals repeated hypoxia events over the past 500 yr in the northern Adriatic Sea. In Geology, 2017, vol. 45, no. 4, p. 363-366.

[50] TOMAŠOVÝCH, Adam – JABLONSKI, David. Decoupling of latitudinal gradients in species and genus geographic range size: a signature of clade range expansion. In Global Ecology and Biogeography, 2017, vol. 26, no. 3, p. 288-303.

[49] TOMAŠOVÝCH, Adam – SCHLÖGL, Ján – BIROŇ, Adrián – HUDÁČKOVÁ, Natália – MIKUŠ, Tomáš. Taphonomic clock and bathymetric dependence of cephalopod preservation in bathyal, sediment-starved environments. In Palaios, 2017, vol. 32, no. 3, p. 135-152.

[48] ZUSCHIN, Martin – NAWROT, Rafał – HARZHAUSER, Mathias – MANDIC, O. – TOMAŠOVÝCH, Adam. Taxonomic and numerical sufficiency in depth- and salinity-controlled marine paleocommunities. In Paleobiology, 2017, vol. 43, no. 3, p. 463-478.

[47] Tomašových A., Schlögl J., Kaufman D.S., Hudáčková N. 2016. Temporal and bathymetric resolution of nautiloid death assemblages in stratigraphically-condensed oozes (New Caledonia). Terra Nova 28, 271-278.

[46] Tomašových A., Kennedy J.D., Betzner T.J., Bitler Kuehnle N., Edie S., Kim S., Supriya K., White A.E., Rahbek, C., Huang S., Price T.D., and Jablonski D. 2016. Unifying latitudinal gradients in range size and richness across marine and terrestrial systems. Proceedings of the Royal Society B 283, 20153027.

[45] Albano, P.G., Filippova, N., Steger, J., Schmidbaur, H., Tomašových, A., Stachowitsch, M. and Zuschin, M., 2016. Contamination patterns and molluscan and polychaete assemblages in two Persian (Arabian) Gulf oilfields. Marine Ecology in press.

[44] Albano, P.G., Filippova, N., Steger, J., Kaufman, D.S., Tomašových, A., Stachowitsch, M. and Zuschin, M. 2016. Oil platforms in the Persian (Arabian) Gulf: Living and death assemblages reveal no effects. Continental Shelf Research, 121, 21-34.

[43] Albano, P. G., Tomašových, A., Stachowitsch, M., and Zuschin, M. 2016. Taxonomic sufficiency in a live–dead agreement study in a tropical setting. Palaeogeography Palaeoclimatology Palaeoecology 449, 341-348.

[42] Chattopadhyay, D., Zuschin, M. and Tomasovych, A., 2015. How effective are ecological traits against drilling predation? Insights from Recent bivalve assemblages of the northern Red Sea. Palaeogeography, Palaeoclimatology, Palaeoecology 440, 659-670.

[41] Tomašových A., Kidwell S.M., Foygel Barber R. 2016. Inferring skeletal production from time-averaged assemblages: skeletal loss pulls the timing of production pulses towards the modern period. Paleobiology 42, 54-76.

[40] Tomašových A., Jablonski D., Berke S.K., Krug Z.A. And Valentine J.W. 2015. Nonlinear thermal gradients shape broad-scale patterns in geographic range size and can reverse Rapoport’s rule. Global Ecology and Biogeography 24, 257-267

[39] Tomašových A., Dominici S., Zuschin M. and Merle D. 2014. Onshore–offshore gradient in metacommunity turnover emerges only over macroevolutionary time-scales. Proceedings of the Royal Society B 281, 20141533.

[38] Tomašových A., Kidwell S.M., Foygel Barber, R. and Kaufman, D.S. 2014. Long-term accumulation of carbonate shells reflects a 100-fold drop in loss rate. Geology 42, 9, 819-822

[37] Chattopadhyay D., Zuschin M. and Tomašových A. 2014. Effects of a high-risk environment on edge-drilling behavior: inference from Recent bivalves from the Red Sea. Paleobiology 40, 34-49.

[36] Vollstaedt, H., Eisenhauer, A., Wallmann, K., Böhm, F., Fietzke, J., Liebetrau, V., Krabbenhöft, A., Farkaš, J., Tomašových, A., Raddatz, J., Veizer, J. 2014. The Phanerozoic δ88/ 86Sr record of seawater: New constraints on past changes in oceanic carbonate fluxes. Geochimica et Cosmochimica Acta  128, 249-265

[35] Kidwell S.M. and Tomašových A. 2013. Implications of time-averaged death Assemblages for ecology and conservation biology. Annual Reviews of Ecology, Evolution, and Systematics 44: 539-563

[34] Jablonski D., Belanger C.L., Berke S.K., Huang S., Krug A.Z., Roy K., Tomasovych A. and Valentine J.W. 2013. Out of the tropics, but how? Fossils, bridge species, and thermal ranges in the dynamics of the marine latitudinal diversity gradient. Proceedings of the National Academy of Sciences USA 110, 10487-10494.

[33] Hillebrandt A. v., Krystyn, L., Kurschner, W.M., Bonis N.R., Ruhl, M., Richoz, S., Schobben, M.A., Urlichs, M., Bown, P.R., Kment, K., McRoberts, C.A., Simms M. and Tomasovych A. 2013. The global stratotype sections and point (GSSP) for the base of the jurassic system at kuhjoch (Karwendel Mountains, Northern Calcareous Alps, Tyrol, Austria). Episodes 36, 162-198.

[32] Simo V. and Tomašových A. 2013. Trace-fossil assemblages with a new ichnogenus in “spotted” (Fleckenmergel-Fleckenkalk) deposits: a signature of oxygen-limited benthic communities. Geologica Carpathica 64, 355-374.

[31] Berke, S.K., D. Jablonski, A., Z. Krug, K. Roy, A. Tomašových, J. W. Valentine. 2012. Beyond Bergmann’s Rule: size-latitude relationships in marine Bivalvia worldwide. Global Ecology and Biogeography 22, 173-183

[30] Reolid, M., García-García. F., Tomašových, A., Soria, J.M, 2012. Thick brachiopod shell concentrations from prodelta and siliciclastic ramp in a Tortonian Atlantic-Mediterranean Strait (Miocene, Guadix Basin, Southern Spain). Facies 58: 549-571

[29] Tomašových A., and Kidwell, S.M. 2011. Accounting for the effects of biological variability and temporal autocorrelation in assessing the preservation of species abundance. Paleobiology 37:332-354

[28] Tomašových A., and Kidwell, S.M. 2010. Predicting the effects of increasing temporal scale on species composition, diversity, and rank-abundance distributions. Paleobiology 36: 672-695

[27] Tomašových A., and Kidwell, S.M. 2010. The effects of temporal resolution on species turnover and on testing metacommunity models. American Naturalist 175:587-606.

[26] Tomašových A., and Zuschin M. 2009. Variation in brachiopod preservation along a carbonate shelf-basin transect (Red Sea and Gulf of Aden): environmental sensitivity of taphofacies. Palaios 24:697-716

[25] Tomašových A., and Kidwell S.M. 2009. Fidelity of variation in species composition and diversity partitioning by death assemblages: time averaging transfers diversity from beta to alpha levels. Paleobiology 35:97-121.

[24] Tomašových A., and Kidwell S.M. 2009. Preservation of spatial and environmental gradients by death assemblages. Paleobiology 35:122-148.

[23] Schlögl, J., Mangold, C., Tomašových A., and Golej, M. 2009. Early and Middle Callovian ammonites from the Pieniny Klippen Belt (Western Carpathians) in hiatal successions: unique biostratigraphic evidence from sediment-filled fissure deposits. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 252:55-79.

[22] Tomašových A., Carlson, S.J., and LaBarbera M. 2008. Ontogenetic niche shift in the brachiopod Terebratalia transversa: relationship between the loss of rotation ability and allometric growth. Palaeontology 51:1471-1496.

[21] Tomašových A. and J. Schlögl. 2008. Analyzing variations in cephalopod abundances in shell concentrations: the combined effects of production and density-dependent cementation rates. Palaios 23:648-666.

[20] Tomašových A. 2008: Substrate exploitation and resistance to biotic disturbance in the brachiopod Terebratalia transversa and the bivalve Pododesmus macrochisma. Marine Ecology-Progress Series 363: 157-170.

[19] Tomašových A. 2008: Evaluating neutrality and escalation hypothesis in brachiopod communities from shallow, high-productivity habitats. Evolutionary Ecology Research 10: 667-698.

[18] Hautmann, M., Benton, M.J., and Tomašových A. 2008. Catastrophic ocean acidification at the Triassic-Jurassic boundary. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 249: 119-127.

[17] van de Schootbrugge, B., Payne, J.L., Tomašových A., Pross, J., Fiebig, J., Benbrahim, M., Föllmi, K.B., Quan, T.M., 2008. Carbon cycle peturbation and stabilization in the wake of the Triassic-Jurassic boundary mass-extinction event. Geochemistry Geophysics Geosystems, v. 9, art. No Q02408.

[16] Yanes Y., Tomašových A., Kowalewski M., Castillo C., Aguirre J., Alonso M.R., and Ibáñez M. 2008. Taphonomy and compositional fidelity of Quaternary fossil assemblages of terrestrial gastropods from carbonate-rich environments of the Canary Islands. Lethaia 41: 235-256.

[15] Alroy J., Aberhan M., Bottjer D.J., Foote M., Fürsich F.T., Harries P.J., Hendy A.J.W., Holland S.M., Ivany L.C., Kiessling W., Kosnik M.A., Marshall C.R., McGowan A.J., Miller A.I., Olszewski T.D., Patzkowsky M.E., Peters S.E., Villier L., Wagner P.J., Bonuso N., Borkow P.S., Brenneis B., Clapham M.E., Fall L.M., Ferguson C.A., Hanson V.L., Krug A.Z., Layou K.M., Leckey E.H., Nürnberg S., Powers C.M., Sessa J.A., Simpson C., Tomašových A., and Visaggi C.C. 2008. Phanerozoic Trends in the Global Diversity of Marine Invertebrates. Science 5885: 97-100.

[14] Tomašových A. 2008. Composition and distribution of Hettangian brachiopod communities in the West Carpathians: recovery after the end-Triassic mass extinction event. Fossils and Strata 54: 173-181.

[13] Logan A., Tomašových A., Zuschin M. and Grill B. 2008. Recent brachiopods from the Red Sea and Gulf of Aden. Fossils and Strata 54: 299-309.

[12] Tomašových A. and Siblík M. 2007. Evaluating compositional turnover of brachiopod communities during the end-Triassic mass extinction (Northern Calcareous Alps): removal of dominant groups, recovery and community re-assembly. Palaeogeography, Palaeoclimatology, Palaeoecology 244: 170-200.

[11] Tomašových A. 2006. Brachiopod and bivalve ecology in Late Triassic (Alps, Austria): onshore-offshore replacements caused by variations in sediment and nutrient supply. Palaios 21, 344-368.

[10] Tomašových A. 2006: A new Early Jurassic rhynchonellid brachiopod from the Western Tethys and implications for systematics of rhynchonellids from the Triassic-Jurassic boundary. Journal of Paleontology 80, 212-228.

[9] Tomašových A. 2006. Linking taphonomy to community-level abundance: insights into compositional fidelity of the Upper Triassic shell concentrations (Eastern Alps). Palaeogeography, Palaeoclimatology, Palaeoecology 235, 355-381.

[8] Tomašových A., Fürsich F.T., and Wilmsen M. 2006: Preservation of autochthonous shell beds by positive feedback between increased hardpart-input rates and increased sedimentation rates. Journal of Geology 114, 287-312.

[7] Tomašových A., Fürsich F. T., and Olszewski T. D. 2006. Modeling the shelliness and alteration in shell beds: opposing predictions due to variations in hardpart-input rates and burial rates. Paleobiology 32, 278-298.

[6] Tomašových A. and Rothfus T.A. 2005: Differential taphonomy of modern brachiopods (San Juan Islands, Washington State): effect of intrinsic factors on damage and community-level abundance. Lethaia 38, 271-292.

[5] Tomašových A. and Farkaš J. 2005: Cathodoluminescence of Late Triassic terebratulid brachiopods: implications for growth patterns. Palaeogeography, Palaeoclimatology, Palaeoecology  216, 215-233.

[4] Tomašových A. 2004: Postmortem durability and population dynamics affecting the fidelity of size-frequency distributions. Palaios 19, 477-496.

[3] Tomašových A. 2004: Effect of extrinsic factors on biofabric and brachiopod alteration in a shallow intraplatform carbonate setting (Upper Triassic, West Carpathians). Palaios 19, 349-371.

[2] Tomašových A. 2004, Microfacies and depositional environment of an Upper Triassic intra-platform carbonate basin: the Fatric Unit of the West Carpathians (Slovakia). Facies 50, 77-105.

[1] Gazdzicki A., Michalík J. and Tomašových A. 2000, Parafavreina coprolites from the Uppermost Triassic of the Western Carpathians. Geologica Carpathica 51, 245-250.